© Benaki Phytopathological Institute
Scarabaeidae Collection in Goulandris Natural History Museum
85
himself.
We followed the systematics of Ratc-
liffe and Jameson (2004) and Triplehorn and
Johnson (2005).
Catalogue of Coleoptera, Scarabaei-
dae
In the list below, we record the species and the
relevant metadata: the number of specimens
(spm), the place of the collection, and the year
and collector’s name, when available.
Dynastinae (21 species, 105 spm)
Pentodon dispar
(syn.
P. algerinum
sbsp.
dispar)
23 spm,
EGYPT: Alexandria, Victo-
ria?, Mariout, Ramleh, Rond-Point, Roun-
leh, Abou el Chekark, 1917, 1921, 1924, 1929,
1935, 1938 leg. Efflatoun
Pentodon punctatus
1 spm, ITALY: Gerace Ca-
labria, leg. Schuster
Pentodon idiota
1 spm, AUSTRIA: Neusiedi
lake, leg. Schuster
Pentodon bispinosus
1spm, MOROCCO: Casa-
blanca, 1919, leg. Antoine
Pentodon
sp.1 spm, TURKESTAN: Samarkand
Pentodon
sp.1 spm, TURKESTAN: Samarkand
Dyscinetus rugifron
2 spm, ARGENTINA: Bue-
nos Aires
Pachononyx fuscodeneus
2 spm, ARGENTINA:
Rio Salodo
Phyllognathus silenus
12 spm, FRANCE: Tou-
louse; TURKEY: Adana, Cilicie; SPAIN: Tener-
ife; EGYPT: Mariout, Ramleh, Palais; ALGERIA:
Bou-Berak, 1921, 1924, 1935, 1953, leg.V.M.
Duchon, Efflatoun, A. Chobaut, Thery, A.
Carneri
Heteronychus parumpunctatus
16 spm,
EGYPT: Alexandria, 1921, 1922, 1924, 1929
Heteronychus dilatatus
2 spm, ARGENTINA:
Tucuman
Heteronychus
spp.
3 spm, EGYPT: Mariout,
1933
Oryctomorphus maculicollis
1 spm, CHILE
Oryctomorphus
sp
.
2 spm, SOUDAN: FRANCE
Temnorrhynchus baal
21 spm, EGYPT: Alex-
andria, Victoria?, Mazarita, Mandara Cairo,
Rond Point, Nousha, 1919, 1921, 1923, 1924,
1932, 1942, leg. A. Carneri
Phileurus valgus
4 spm, VENEZUELA
Scaptophilus dasipleurus
2 spm, ARGENTINA:
Tucuman
Oryctes rhinoceros
2 spm, INDONESIA: Am-
boine, Iles Moluques, leg. A. Carneri
Oryctes
spp. 5 spm, ARGENTINA: Tucuman
Oryctes
sp. 2 spm, ITALY, 1950, leg. Gentile
Oryctes nasicornis
1 spm
Melolonthinae (11 species, 30 spm)
Europtron gracile
4 spm, ALGERIA: Ghardaia,
leg. A. Chobaut
Haplidia nitidula
7 spm
Hoplia coerulea
1 spm, FRANCE: Prats de
Mollo, leg. Waegner
Hoplia dubia
5 spm, ITALY: Macarese, Rome,
1919, leg. P. Crozet
Hoplia africanum
4 spm, MOROCCO: Casa-
blanca, leg. Antoine
Hoplia farinosa
2 spm, AUSTRIA: Graz;
FRANCE: Besançon
Hoplia philanthus
1 spm
Hoplia graminicola
1 spm
Hoplia praticola
1 spm, ALGERIA: Ain Sefra,
1923, leg. A. Chobaut
Hoplia pubicollis
1 spm, CORSICA: Ajaccio
Hoplia
spp. 3 spm, ALGERIA: Bou-berak; MO-
ROCCO: Casablanca, leg. A. Chobaut, An-
toine
Rutelinae (29 species, 71 spm)
Anisoplia flavipennis
1 spm
Anisoplia segetum var. straminea
(syn.
Chaetopteroplia segetum straminae
)
1 spm,
TURKEY: Konya, 1899, leg. Korh
Anisoplia segetum
(syn.
Chaetopteroplia sege-
tum)
1 spm, IRAN: Kopet-Dagh, Descht, 1902,
leg. F. Hauser
Anisoplia pallidipennis
2 spm, ALGERIA: Ain
Sefra, 1923, leg. A. Chobaut
Anisoplia cyathigera
1 spm, BOSNIA: Sanski-
most
Anisoplia biguttata
1 spm, INDONESIA: Java
Breanger Tjigembong,
leg. Corporaai
Anisoplia lata
2 spm,
AUSTRIA: Graz
,
CROA-
TIA: Pola Istria
Anisoplia leucaspis
1 spm, IRAN: Kerman-
schah, 1909, leg. F. Hauser
Anisoplia agricola
1 spm, GERMANY: Offen-
bach
Anisoplia
sp
.
1 spm,
CROATIA: Velezgb,
leg.
Krauss