Travlos
        
        
          et al.
        
        
          110
        
        
          Science Department at the Benaki Phy-
        
        
          topathological Institute. These characters
        
        
          can provide a safe recognition of the spe-
        
        
          cies. In actual measurements of spikelets
        
        
          in the field the mean length of the awns,
        
        
          for example, was 5.514 cm for
        
        
          A. sterilis
        
        
          and 3.625 cm for
        
        
          A. fatua
        
        
          . Also, the mean
        
        
          length of the florets was 2.392 and 1.85
        
        
          cm for
        
        
          A. sterilis
        
        
          and
        
        
          A. fatua
        
        
          , respective-
        
        
          ly. Moreover, a difference was observed in
        
        
          the time of maturation,
        
        
          A. fatua
        
        
          maturing
        
        
          about 10-15 days later than
        
        
          A. sterilis,
        
        
          and
        
        
          this
        
        
          is likely associated with different tem-
        
        
          perature requirements for germination;
        
        
          optimum germination temperatures for
        
        
          A.
        
        
          sterilis
        
        
          are reported to be lower than those
        
        
          for
        
        
          A. fatua
        
        
          (6, 17).
        
        
          It is worthy to mention that the above
        
        
          characters can provide a safe recognition
        
        
          of the species if used on the seeds as they
        
        
          are shedding. However, processing of the
        
        
          seeds after shedding, which may cause
        
        
          breaking of awns, rubbing off the hairs,
        
        
          breaking of florets or bases etc. is expect-
        
        
          ed to reduce the diagnostic value of these
        
        
          characters.
        
        
          The wild oat accessions from the Viotia
        
        
          region, when grown under the same con-
        
        
          ditions in pots in the greenhouse, exhibit-
        
        
          ed a significant variation in growth habit
        
        
          and other parameters of seedling devel-
        
        
          opment (Table 3). Variability was wide in
        
        
          the
        
        
          A. sterilis
        
        
          accessions but very limited in
        
        
          the
        
        
          A. fatua
        
        
          accessions.
        
        
          The most important variation found
        
        
          was related to the growth habit of the
        
        
          A.
        
        
          sterilis
        
        
          accessions, in some of which the
        
        
          seedlings had an erect growth pattern, in
        
        
          some others a semi-erect pattern and in
        
        
          others a prostrate growth pattern (Figure
        
        
          2). In all
        
        
          A. fatua
        
        
          accessions from Viotia the
        
        
          seedlings exhibited only the erect growth
        
        
          habit. The peculiar prostrate pattern of
        
        
          growth for some
        
        
          A. sterilis
        
        
          accessions has
        
        
          also been observed by other researchers
        
        
          in Greece (3) as well as in other countries
        
        
          (12, 16).
        
        
          Significant differences among wild
        
        
          oat accessions were also observed when
        
        
          plant height and tillering were compared
        
        
          (Table 3). Tillering, measured as the num-
        
        
          ber of tillers per plant, seems to correlate
        
        
          negatively with plant height and posi-
        
        
          tively with the growth habit, being high-
        
        
          er in plants of semi-erect and highest in
        
        
          plants of the prostrate growth pattern, in
        
        
          the
        
        
          A. sterilis
        
        
          accessions. In the
        
        
          A. fatua
        
        
          ac-
        
        
          cessions, in which only the erect growth
        
        
          pattern was observed, tillering was mini-
        
        
          
            Figure 2.
          
        
        
          Growth habit of
        
        
          A. sterilis
        
        
          accessions from the Vi-
        
        
          otia region in Greece. Erect (A), Semi-erect (B) and Prostrate
        
        
          (C) seedling growth pattern .
        
        
          
            Table 2.
          
        
        
          Mature seed characters discriminating
        
        
          A. fatua
        
        
          from
        
        
          A. sterilis
        
        
          in the field
        
        
          
            .
          
        
        
          Character
        
        
          Avena sterilis
        
        
          Avena fatua
        
        
          Maturity
        
        
          Earlier maturing
        
        
          Later maturing
        
        
          Awn length (cm)
        
        
          3-7
        
        
          3-4
        
        
          Awn attachment
        
        
          Below middle of floret
        
        
          Above middle of floret
        
        
          Floret length (cm)
        
        
          1.7-3.0
        
        
          1.4-2.1
        
        
          Basal scar shape
        
        
          Large, elongated
        
        
          Absent or small round
        
        
          Hair covering
        
        
          50-80% of floret surface
        
        
          30-60% of floret surface